Trees, fairly large in natural vegetation; clonal trees small, 4-7 m high and usually with a spreading habit. Leaves alternate, paripinnate, up to 6-jugate; leaflets ovate to obovate, 5-28 cm x 2-10.5 cm, usually horizontal, above glabrous or sometimes slightly hairy on the midrib, beneath variably hairy, domatia common or absent, apex truncate to acuminate, nerves slightly to strongly curving, veins scalariform to coarsely reticulate. Inflorescence pseudo-terminal to usually terminal. Flower either male(only stamens well developed) or hermaphrodite and either effectively female (stamens small, anther not dehiscing) or male (stigma not opening), actinomorphic, whitish, yellowish or greenish; sepals 4-5, nearly free to more than halfway connate, 0.7-2.1 mm long. Petals usually absent, sometimes up to 4 reduced ones, not exceeding 1.6 mm. Disc complete, hairy or glabrous. Stamens (4)- 5-8 (-9), exserted in males; filaments has dense long hairs at least in the basal part; anther dehiscing latero-introrse, lengthwise. Pistil 2- or rarely 3-merous, densely hairy, well developed in hermaphrodite flowers; ovary warty; style well-developed; stigma spreading to finally recoiled. Fruits ellipsoid to subglobular, up to 6 cm x 3.5 cm, usually only 1 locule, yellow to purplish red, hardly stalked, apparently often at least in the apical part finely dehiscing, glabrous, usually densely set with filiform, curved, 0.5-2 cm long appendages; seed covered by a thick, juicy, white to yellow, translucent sarcotesta.